M = marine species

SUPPLEMENTARY TABLE 1. Specimens examined, habitat and measured values of
parameters: M = marine species; F = freshwater species; L = land species; SUM =
Shizuoka University; SHIN = Shinshu University; BPBM = Bishop Museum; SPMN =
Network for Shizuoka Prefecture Museum of Natural History; UMUT = University
Museum, University of Tokyo.
Species
Orthogastropoda
Vetigastropoda
Haliotoidea
Haliotidae
Haliotis gigantea
Haliotis diversicolor aquatilis
Pleurotomarioidea
Pleurotomariodae
Perotrochus hirasei
Perotrochus africanus westralis
Trocoidea
Trochidae
Chlorostoma xanthostigma
Chlorostoma xanthostigma
Granata lyrata
Euchelus atrata
Cittarium pica
Lischkeia alwinae
Ginebis argenteonitens form convexiuscula
Trochus erythraeus
Trochus maculatus
Tectus conus
Tectus triserialis
Tegula regina
Bankivia fasciata
Monodonta turbinata
Fossarina petterdi
Stomatia phymotis
Calliostoma granulatum
Umbonium giganteum
Repository Habitat
W
model parameters
T
D
!
"
functional parameters
1/#
E
SPMN
SUM
M
M
16.815
11.907
0.696
0.449
-0.184
-0.241
1.136
1.102
-0.138
0.088
0.203
0.332
0.809
0.792
SPMN
SPMN
M
M
1.680
1.929
1.256
1.146
0.016
0.003
0.902
0.900
0.056
-0.100
0.181
0.220
0.822
0.811
UMUT
UMUT
UMUT
UMUT
UMUT
SPMN
SPMN
SPMN
UMUT
SPMN
SUM
SPMN
UMUT
UMUT
UMUT
UMUT
SPMN
UMUT
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
2.127
1.893
2.826
2.296
2.142
1.678
1.658
1.611
1.441
1.465
1.402
1.693
1.425
1.857
4.377
5.293
1.746
1.910
1.192
1.308
1.261
1.437
1.417
1.354
1.934
1.434
1.917
2.092
2.054
1.626
4.146
1.560
0.962
2.259
1.512
0.960
-0.655
-0.231
-0.321
-0.175
-0.248
-0.165
-0.067
-0.160
-0.274
-0.151
0.059
0.019
-0.078
-0.181
0.059
-0.091
-0.081
0.172
1.086
1.102
0.798
0.883
0.965
1.030
0.544
1.103
1.183
1.067
1.012
1.309
0.420
0.922
0.733
0.483
0.797
0.778
-0.567
-0.166
0.286
-0.409
-0.206
-0.408
-0.174
-0.380
-0.131
-0.279
-0.147
0.517
-0.112
-0.462
-0.630
-0.415
-0.429
-0.192
0.184
0.193
0.199
0.195
0.221
0.235
0.185
0.222
0.183
0.179
0.200
0.138
0.147
0.238
0.167
0.135
0.199
0.184
0.558
0.783
0.781
0.802
0.782
0.783
0.817
0.789
0.755
0.800
0.814
0.799
0.810
0.799
0.809
0.809
0.806
0.809
Species
Turbinoidea
Turbinidae
Liotina peronii
Cookia sulcata
Astraea undosa
Angaria delphinus
Angaria delphinus
Angaria neglecta
Turbo cornutus
Turbo petholatus
Astralium haematragum
Astralium haematragum
Guildfordia triumphans
Phasianella australis
Phasianella australis
Neritimorpha
Cyclonertimorpha
Neritoidea
Neritidae
Nerita albicilla
Nerita costata
Nerita exuvia
Nerita japonica
Neritina cornucopia
Apogastropoda
Caenogastropoda
Architaenioglossa
Ampullaroidea
Ampullariidae
Lanistes olivaceus
Repository Habitat
W
model parameters
T
D
!
"
functional parameters
1/#
E
UMUT
UMUT
SPMN
UMUT
SPMN
UMUT
SUM
SPMN
SUM
SUM
SUM
UMUT
SPMN
M
M
M
M
M
M
M
M
M
M
M
M
M
2.344
1.734
1.878
2.494
2.388
2.344
2.039
2.292
1.542
1.833
2.102
1.724
1.689
1.060
1.271
1.413
0.998
0.679
0.998
1.463
1.331
1.899
1.357
0.888
2.455
2.805
-0.331
-0.141
0.082
0.047
0.087
0.002
-0.282
-0.051
-0.013
0.016
0.157
-0.145
-0.041
0.441
0.897
1.041
0.535
0.303
0.656
0.632
0.516
1.070
1.015
0.684
0.411
0.274
-0.319
0.060
-0.483
-0.291
-0.201
-0.466
-0.375
-0.378
-0.244
-0.129
-0.182
-0.417
-0.030
0.197
0.174
0.174
0.211
0.161
0.164
0.237
0.229
0.193
0.189
0.181
0.190
0.153
0.764
0.799
0.808
0.809
0.817
0.809
0.786
0.808
0.807
0.808
0.809
0.804
0.809
SUM
SPMN
SPMN
UMUT
UMUT
M
M
M
M
M
19.372
4.819
8.479
3.513
9.654
0.189
1.013
0.873
1.044
0.263
-0.021
-0.283
-0.048
-0.064
-0.031
-0.006
0.634
0.345
0.696
0.066
-0.211
-0.758
-0.209
-0.543
-0.153
0.318
0.232
0.212
0.160
0.235
0.810
0.791
0.809
0.808
0.810
UMUT
F
1.763
2.104
-0.059
0.358
0.296
0.150
0.807
Species
Pomacea canaliculata
Pomacea depressa
Pomacea glauca
Cyclophroidea
Cyclophoridae
Cyclopharus miyakoensis
Cyclophorus appendiculayus
Cyclophorus turgidus turgidus
Cyclophorus zebrinus
Leptopoma nitidum
Pterocyclus perrieri
Pupinidae
Pupinella rufa
Orcula dolium
Viviparoidea
Viviparidae
Cipangopaludina chinensis laeta
Heterogen longispira
Heterogen longispira
Melantho poderosa
Sinotaia quadrata histrica
Tulotoma angulata
Viviparus contectoides
Sorbeoconcha
Cerithoidea
Certhiidae
Cerithium nodulosus
Rhinoclavis fasciata
Rhinoclavis vertagus
Turritellidae
SPMN
UMUT
UMUT
F
F
F
W
2.183
2.487
2.169
model parameters
T
D
0.563
0.065
1.064
0.011
1.124
0.050
UMUT
SPMN
SPMN
SPMN
UMUT
SHIN
L
L
L
L
L
L
2.029
2.339
2.027
2.062
1.974
2.441
1.096
0.764
1.033
0.987
1.338
0.404
0.181
0.239
0.135
0.007
0.088
0.241
0.560
0.332
0.561
0.657
0.569
0.795
-0.370
-0.268
-0.033
-0.088
-0.260
0.046
0.184
0.152
0.164
0.188
0.191
0.145
0.810
0.810
0.812
0.817
0.810
0.817
UMUT
UMUT
L
L
1.328
1.187
4.363
4.793
-0.149
0.215
-0.133
0.151
-0.664
-1.052
0.141
0.126
0.785
0.802
UMUT
UMUT
SPMN
UMUT
SPMN
UMUT
SPMN
F
F
F
F
F
F
F
1.969
1.540
1.535
1.763
1.700
1.862
1.924
1.519
2.430
2.412
2.120
2.119
1.586
1.644
-0.057
0.037
0.047
-0.041
0.000
-0.064
0.015
0.526
0.609
0.549
0.367
0.580
0.770
0.364
-0.102
-0.396
-0.213
-0.182
-0.327
-0.533
-0.214
0.192
0.186
0.171
0.173
0.196
0.199
0.175
0.811
0.809
0.809
0.809
0.809
0.806
0.809
SPMN
SPMN
SUM
M
M
M
1.252
1.136
1.250
4.863
9.584
4.722
0.053
-0.432
-0.143
-0.333
-0.025
0.088
-0.369
-0.109
-0.103
0.130
0.108
0.138
0.808
0.710
0.811
Repository Habitat
!
0.286
0.218
0.253
"
-0.280
-0.105
-0.154
functional parameters
1/#
E
0.158
0.811
0.165
0.811
0.162
0.815
Species
Turritella rubescens
Turritella fortilirata
Modulidae
Modulus candidus
Planaxidae
Planaxis sulcatus
Quoyia decollata
Potamididae
Certhidea obtusa
Tympanotonus fuscatus
Telebralia palustris
Pleuroceridae
Anculosa praerosa
Semisulcospira niponica
Thiaridae
Melanoides tuberculatus
Semisulcospira reticulata
Faunus ater
Hypsogastropoda
Littorinimorpha
Littorinoidea
Littorinidae
Echininus cumingii spinulosus
Lacuna vincta
Littorina littorea
Littorina undulata
Littorina zebra
Tectarius coronatus
Truncatelloidea
Rissoidae
SPMN
SUM
M
M
W
1.169
1.205
model parameters
T
D
!
9.464
0.023
0.358
6.865
-0.091
0.492
UMUT
M
2.402
0.809
-0.201
0.625
-0.140
0.192
0.791
UMUT
UMUT
M
M
1.648
1.535
2.487
2.997
-0.095
-0.183
0.495
0.452
-0.032
-0.554
0.175
0.183
0.811
0.797
UMUT
UMUT
SUM
M
M
M
1.264
1.169
1.199
4.570
6.431
4.763
-0.069
-0.138
0.018
-0.174
-0.319
-0.228
-0.393
0.414
-0.187
0.143
0.135
0.149
0.800
0.797
0.823
UMUT
UMUT
F
F
3.522
1.414
1.280
3.829
-0.116
-0.063
0.144
0.296
-0.217
-0.179
0.193
0.147
0.806
0.809
UMUT
SUM
SPMN
F
F
F
1.337
1.192
1.235
5.027
6.254
6.318
-0.074
-0.214
-0.176
0.366
0.656
-0.160
0.173
-0.369
-0.181
0.128
0.137
0.123
0.809
0.810
0.786
UMUT
UMUT
UMUT
UMUT
SPMN
SPMN
M
M
M
M
M
M
1.740
2.475
2.101
1.794
2.317
1.612
1.551
2.052
1.656
2.419
1.498
1.937
0.021
-0.182
-0.148
0.008
-0.004
-0.180
0.787
0.549
0.597
0.418
0.768
0.692
-0.058
-0.067
-0.238
0.122
-0.353
-0.472
0.196
0.183
0.234
0.158
0.162
0.218
0.809
0.810
0.806
0.809
0.808
0.806
Repository Habitat
"
-0.479
-0.446
functional parameters
1/#
E
0.101
0.809
0.126
0.806
Species
Rissona marginata
Pomatiopsidae
Oncomelania nosophora
Assimineidae
Assiminea grayana
Stromboidea
Strombidae
Lambis chiragra
Lambis lambis
Strombus aurisdianae
Strombus dentatus
Strombus pipus
Strombus urceas
Strombus vomer
Tibia fusus
Tibia insulaechorab
Seraphidae
Terebellum terebellum terebellum
Hipponix conica
Aporrhaidae
Aporrhais pespelecani
Aporrhais occidentalis
Struthiolariidae
Struthiolaria papulosa
Vanikoloidea
Vanikoridae
Vanikoro acuta
Capuloidea
Capulidae
Capulus ungaricus
UMUT
M
W
1.714
model parameters
T
D
!
2.597
-0.118
0.547
UMUT
F
1.331
5.172
0.286
0.044
-0.244
0.105
0.810
UMUT
M
1.786
2.518
-0.316
0.435
0.000
0.168
0.783
SUM
UMUT
UMUT
UMUT
SPMN
SUM
SPMN
SPMN
SUM
M
M
M
M
M
M
M
M
M
1.733
3.888
1.641
1.564
1.824
1.521
1.572
1.233
1.313
1.976
1.860
1.741
2.686
1.282
2.003
2.054
6.785
4.348
-0.138
-0.421
-0.217
-0.197
0.108
0.036
-0.100
-0.127
0.075
0.036
0.026
-0.018
0.189
0.192
0.143
-0.106
0.271
-0.127
-0.143
-0.928
0.195
0.159
-0.494
0.284
0.363
-0.023
-0.258
0.150
0.169
0.152
0.144
0.162
0.142
0.142
0.119
0.131
0.808
0.749
0.789
0.799
0.819
0.824
0.807
0.807
0.809
SPMN
UMUT
M
M
2.216
46.615
5.396
0.285
-0.399
0.099
-0.083
-0.177
0.000
-0.448
0.103
0.147
0.772
0.810
SPMN
UMUT
M
M
1.436
1.416
3.177
3.622
-0.010
0.004
-0.280
-0.248
-0.241
-0.173
0.138
0.133
0.806
0.807
UMUT
M
1.630
2.563
-0.317
0.288
-0.269
0.160
0.771
UMUT
M
3.168
1.125
-0.101
0.597
-0.266
0.202
0.808
SPMN
M
135.702
0.085
0.044
0.105
-0.695
0.202
0.809
Repository Habitat
"
-0.208
functional parameters
1/#
E
0.194
0.810
Species
Trichamathina nobilis
Trichotropis bicarinata
Calyptraeoidea
Calyptraeidae
Crepidula fornicata
Xenophoroidea
Xenophoridae
Stellaria exutus
Stellaria solaris
Xenophora pallidula
Cypraeoidea
Cypraeidae
Cypraea mauritiana
Cypraea tigris
Ovulidae
Volva volva volva
Ovula ovum
Velutinoidea
Triviidae
Erato voluta
Naticoidea
Naticidae
Cryptonatica andoi
Euspira fortunei
Glossaulax bicolor
Glossaulax didyma
Glossaulax vesicalis
Lunatia lewisi
Polinices catena
Polinices mammilla
UMUT
UMUT
M
M
W
12.868
2.731
model parameters
T
D
!
0.212
0.159
0.129
0.932
-0.018
0.442
UMUT
M
19.411
0.260
-0.162
0.186
-0.484
0.317
0.805
SUM
SPMN
UMUT
M
M
M
2.095
1.932
1.693
0.815
0.947
1.122
0.127
0.044
-0.040
0.825
1.333
1.232
-0.157
0.507
0.165
0.194
0.136
0.183
0.810
0.807
0.812
UMUT
UMUT
M
M
6.169
2.486
1.613
0.905
-0.057
-0.294
0.213
0.233
-0.149
-0.183
0.167
0.171
0.809
0.774
UMUT
UMUT
M
M
1.215
2.235
3.080
0.764
-0.336
-0.195
0.026
-0.020
-0.109
0.061
0.146
0.143
0.734
0.786
UMUT
M
2.271
1.710
-0.175
0.009
-0.220
0.153
0.794
UMUT
SPMN
UMUT
SUM
SPMN
SPMN
UMUT
SPMN
M
M
M
M
M
M
M
M
2.679
1.929
3.493
3.317
6.108
2.529
2.247
4.291
1.160
1.682
0.869
1.047
1.114
0.993
1.312
1.497
0.046
-0.079
0.044
-0.099
-0.116
-0.096
0.081
-0.095
0.525
0.596
0.713
0.812
0.723
0.665
0.534
0.413
-0.240
-0.179
-0.485
0.123
-0.169
0.047
-0.596
-0.237
0.220
0.209
0.177
0.248
0.181
0.198
0.163
0.169
0.809
0.808
0.809
0.807
0.809
0.814
0.809
0.809
Repository Habitat
"
-0.458
-0.272
functional parameters
1/#
E
0.154
0.810
0.211
0.809
Species
Sinum javanicum
Risooidea
Bithyniidae
Bithynia tentaculata
Parafossarulus manchouricus
Parafossarulus manchouricus
Truncatellidae
Truncatella marginata
Tonnoidea
Bursidae
Bufonaria rana
Bursa elegans
Cassidae
Casmaria ponderosa ponderosa
Casmaria ponderosa ponderosa
Cassis nana
Cassis vibex
Cypraecassis coarctata
Cypraecassis rufa
Galeodea rugosa
Galeodea rugosa
Phalium flammiferum
Semicassis bisulcata pila
Semicassis pyrum
Ficidae
Ficus ventricosa
Ficus subintermedia
Tonnidae
Eudolium lineatum
Malea pomum
UMUT
M
W
6.912
model parameters
T
D
!
0.979
-0.235
0.792
UMUT
UMUT
UMUT
F
F
F
1.967
1.521
1.582
2.010
2.704
3.144
0.036
0.053
0.031
0.410
0.690
0.734
-0.375
-0.461
-0.185
0.188
0.171
0.152
0.809
0.809
0.809
UMUT
L
1.216
6.935
-0.016
0.349
0.162
0.118
0.809
UMUT
SUM
M
M
1.812
1.643
1.625
1.942
-0.311
-0.106
0.228
0.220
-0.042
-0.378
0.154
0.168
0.773
0.815
SPMN
SPMN
SPMN
UMUT
SPMN
SPMN
SPMN
SPMN
SUM
SUM
SPMN
M
M
M
M
M
M
M
M
M
M
M
2.018
1.958
2.636
1.989
1.947
1.461
2.021
1.954
1.831
1.889
2.094
1.945
1.768
0.484
1.838
0.979
1.899
1.177
1.486
1.319
1.576
1.122
0.031
-0.398
0.036
-0.369
-0.228
-0.388
0.076
0.009
-0.309
-0.454
-0.162
0.221
0.102
0.119
0.181
0.095
0.118
0.159
0.152
0.247
0.095
0.246
-0.608
-0.259
-0.843
-0.570
-0.848
-0.596
-0.481
-0.548
-0.797
-0.250
-0.702
0.176
0.172
0.167
0.173
0.185
0.151
0.166
0.171
0.182
0.153
0.198
0.809
0.739
0.806
0.738
0.773
0.696
0.816
0.809
0.738
0.716
0.796
UMUT
SUM
M
M
3.107
1.989
1.057
1.267
-0.330
-0.016
0.030
0.096
-0.238
0.015
0.168
0.163
0.764
0.819
SPMN
SPMN
M
M
1.955
2.120
1.663
0.525
0.166
0.087
0.311
0.145
-0.490
-0.316
0.169
0.174
0.810
0.809
Repository Habitat
"
-0.193
functional parameters
1/#
E
0.191
0.808
Species
Tonna perdix
Tonna luteostoma
Ranellidae
Biplex perca
Charonia lampas macilenta
Charonia variegate
Cymatium gutturnium
Cymatium parthenopeum
Gelagna succincta
Linatella cingulata
Personidae
Distorsio anus
Ptenoglossa
Triphoroidea
Cerithiopsidae
Cerithiopsis emersonii
Triphoridae
Marshallora adversa
Janthinoidea
Janthinidae
Janthina janthina
Janthina janthina
Janthina umbilicata
Epitoniidae
Cirsotrema varicosum
Epitonium scalare
Gyroscala lamellosa
Eulimoidea
Eulimidae
Melanella major
UMUT
SUM
M
M
W
2.694
2.251
model parameters
T
D
!
1.505
-0.314
0.435
1.134
-0.371
0.347
SPMN
UMUT
SPMN
SPMN
SUM
UMUT
SPMN
M
M
M
M
M
M
M
1.485
1.520
1.522
1.849
1.629
1.729
1.885
3.710
2.532
3.165
1.334
2.164
1.813
1.548
-0.083
-0.077
-0.362
0.161
-0.125
-0.246
0.151
0.052
0.173
0.136
0.047
0.241
0.395
0.076
-0.827
-0.933
-0.515
-1.400
-1.304
-0.305
-0.481
0.142
0.168
0.146
0.140
0.171
0.178
0.150
0.789
0.790
0.731
0.769
0.760
0.790
0.810
UMUT
M
1.391
1.852
-0.327
-0.002
-0.873
0.153
0.688
UMUT
M
1.102
7.802
-0.320
0.165
-0.971
0.119
0.648
UMUT
M
1.112
8.228
-0.067
0.706
-0.505
0.125
0.819
UMUT
SPMN
UMUT
M
M
M
3.082
2.922
2.898
1.183
1.107
1.432
-0.009
0.084
-0.203
0.588
0.587
0.203
-0.041
-0.282
-0.443
0.233
0.159
0.213
0.808
0.809
0.797
SPMN
UMUT
UMUT
M
M
M
1.343
1.624
1.463
4.316
2.333
3.205
-0.029
0.036
-0.087
0.367
0.228
0.268
-0.427
-0.379
-0.227
0.145
0.158
0.156
0.808
0.809
0.808
UMUT
M
1.213
5.688
-0.083
0.207
-0.255
0.134
0.818
Repository Habitat
"
-0.342
-0.457
functional parameters
1/#
E
0.234
0.785
0.191
0.745
Species
Scalenostoma carinata
Neogastropoda
Muricoidea
Muricidae
Acanthina monodon
Bolinus cornutus
Ceratostoma burnetti
Chicoreus banksii
Chicoreus cornucervi
Chicoreus torrefactus
Concholepas concholepas
Coralliophila neritoides
Cronia margariticola
Drupa ricinus ricinus
Haustellum haustellum
Homalocantha scorpio
Mancinella echinata
Morula granulata
Murex pecten pecten
Nassa francolina
Neothais orbita
Ocenebra interfossa
Phyllonotus erthrostomus
Pterynotus elongatus
Rapa bubiformis
Rapa rapa
Rapana venosa
Siratus alabaster
Tais armigera
Thais bronni
UMUT
M
W
1.168
model parameters
T
D
!
5.564
0.114
-0.323
UMUT
SPMN
UMUT
SPMN
SPMN
SPMN
UMUT
UMUT
UMUT
SPMN
SPMN
SPMN
SPMN
UMUT
SPMN
UMUT
SPMN
UMUT
SPMN
SPMN
UMUT
SPMN
SUM
SPMN
SPMN
UMUT
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
M
2.857
2.564
1.914
1.580
1.775
1.542
9.393
3.033
1.552
1.895
2.318
2.131
1.805
1.930
1.838
2.295
2.347
1.722
1.801
1.294
2.129
2.158
2.238
1.714
1.716
1.907
1.167
0.902
1.595
2.233
1.810
2.239
0.591
1.146
2.128
1.415
1.169
2.297
1.740
1.543
1.944
2.222
1.580
2.791
1.255
5.876
1.003
0.912
1.488
2.497
1.684
1.551
Repository Habitat
-0.056
-0.091
0.043
0.337
0.241
0.111
-0.136
-0.047
0.280
-0.091
0.170
0.090
0.001
-0.087
0.379
-0.115
-0.139
-0.334
-0.032
0.269
-0.128
-0.069
-0.200
0.017
-0.088
-0.065
0.359
0.232
0.317
0.110
0.204
0.220
-0.061
0.537
0.253
0.456
-0.270
-0.026
0.431
0.490
0.211
0.139
0.350
0.256
0.293
0.085
0.050
0.065
0.296
0.227
0.327
0.374
"
-0.018
-0.564
-0.712
-0.258
-0.436
-0.822
-0.581
-0.767
-0.482
-0.863
-1.357
-0.331
-0.721
-0.631
-0.990
-0.738
-0.218
-0.331
-0.799
-0.879
-1.055
-0.098
-0.671
-0.248
-0.470
-0.293
-0.601
functional parameters
1/#
E
0.143
0.810
0.187
0.215
0.170
0.125
0.158
0.157
0.245
0.156
0.149
0.171
0.128
0.143
0.198
0.193
0.112
0.163
0.209
0.173
0.191
0.096
0.151
0.177
0.192
0.160
0.174
0.200
0.808
0.803
0.809
0.810
0.809
0.809
0.796
0.808
0.809
0.758
0.810
0.808
0.808
0.784
0.810
0.806
0.805
0.733
0.802
0.806
0.805
0.804
0.805
0.809
0.818
0.807
Species
Thais clavigera
Thais haemastoma
Thais kiosquiformis
Trophon geversianus
Urosalpinx cinerea
Vexilla vexillum
Turbinellidae
Columbarium harrisium
Columbarium pagoda pagoda
Vasum ceramicum
Vasum turbinellum
Columbellidae
Amphissa columbiana
Colubraria cumingii
Parametaria philippinarum
Pyrene splendidula
Strombina lanceolata
Nassariidae
Bullia gradata
Nassarius arcularia
Zeuxis margritifera
Buccinidae
Balylonia japonica
Balylonia japonica
Balylonia japonica
Buccinum osagawai
Cominella lineolata
Fusinus dupetitthouarsi
Japelion pericochlion
Kelletia lischkei
SUM
SPMN
UMUT
UMUT
UMUT
UMUT
M
M
M
M
M
M
W
1.962
2.050
1.732
2.261
1.777
2.083
model parameters
T
D
1.651
-0.150
1.801
0.149
1.846
-0.004
1.417
-0.036
2.250
-0.016
1.705
-0.102
SPMN
UMUT
UMUT
UMUT
M
M
M
M
1.419
1.475
1.522
1.753
2.529
2.296
1.961
1.477
0.363
0.122
-0.206
-0.127
0.215
0.201
0.397
0.362
-0.682
-0.395
0.110
-0.932
0.134
0.151
0.151
0.200
0.810
0.810
0.791
0.789
UMUT
SPMN
UMUT
UMUT
UMUT
M
M
M
M
M
1.554
1.301
1.600
1.374
1.448
2.581
3.536
1.802
2.591
3.092
-0.158
-0.083
-0.317
-0.087
0.097
0.296
0.137
0.075
-0.138
0.195
-0.447
-1.138
-0.190
-0.252
-0.759
0.172
0.154
0.150
0.151
0.147
0.809
0.781
0.761
0.815
0.808
SPMN
UMUT
UMUT
M
M
M
1.640
1.739
1.438
2.687
1.592
3.070
-0.196
-0.230
0.069
0.177
0.308
0.279
-0.114
-1.457
-0.817
0.152
0.158
0.154
0.795
0.711
0.805
UMUT
UMUT
UMUT
SPMN
UMUT
SPMN
SUM
UMUT
M
M
M
M
M
M
M
M
1.836
1.849
1.827
1.690
1.622
1.380
1.484
1.609
2.094
2.099
2.089
2.534
2.345
3.878
3.981
2.196
0.083
0.086
0.158
-0.048
-0.253
-0.446
-0.100
-0.359
0.431
0.545
0.567
0.359
0.397
0.097
0.262
0.308
0.020
-0.478
-0.246
-0.428
-0.288
0.602
0.317
-0.119
0.158
0.161
0.179
0.175
0.174
0.135
0.131
0.155
0.809
0.809
0.810
0.808
0.793
0.733
0.805
0.759
Repository Habitat
!
0.286
0.254
0.450
0.174
0.288
0.103
"
-0.704
-0.623
-0.865
-0.477
-0.546
0.226
functional parameters
1/#
E
0.200
0.796
0.173
0.809
0.180
0.799
0.181
0.808
0.175
0.808
0.143
0.803
Species
Neptunea antiqua
Neptunea constricta
Neptunea lyrata decemcostata
Northia northiae
Penion maxima
Pleuroploca princeps
Siphonalia fusoides
Solenosteria fusiformis
Volutharpa ampullacea perryi
Melongenidae
Busycom spiratum
Hemifusus tuba
Melongena melongena
Melongena patula
Pugilina cochlidium
Fasciolariidae
Fasciolaria tulipa
Fusinus forceps
Fusinus nicobaricus
Fusinus nicobaricus
Latirus nodatus
Opeatostoma pseudodon
Peristernia incarnata
Pleuroploca glabra
Volutidae
Amoria undulata
Cymbium olla
Ericusa sericatum
Fulgoraria clara
Lyria planicostata
UMUT
SUM
UMUT
SPMN
SPMN
SPMN
SPMN
UMUT
UMUT
M
M
M
M
M
M
M
M
M
W
1.710
1.622
2.069
1.489
1.606
1.592
1.709
1.614
2.482
model parameters
T
D
2.010
-0.154
2.488
-0.102
1.658
-0.018
3.298
-0.401
2.004
-0.177
2.571
-0.070
2.350
0.165
2.015
-0.035
1.404
-0.290
SPMN
UMUT
SPMN
UMUT
SPMN
M
M
M
M
M
2.254
1.739
1.860
2.316
1.835
0.969
2.076
1.682
1.033
1.324
-0.025
-0.171
-0.030
-0.250
-0.020
0.072
0.187
0.125
0.207
0.127
-0.286
-0.262
-0.079
-0.133
0.045
0.160
0.166
0.154
0.164
0.149
0.817
0.809
0.812
0.785
0.820
SPMN
UMUT
UMUT
SPMN
UMUT
UMUT
UMUT
UMUT
M
M
M
M
M
M
M
M
1.870
1.392
1.422
1.398
1.473
1.735
1.335
1.501
2.242
3.338
2.935
3.198
2.789
1.896
3.297
3.010
-0.133
0.151
0.090
-0.117
-0.134
-0.143
-0.465
-0.227
0.064
0.196
0.300
0.137
0.104
0.184
0.503
0.128
-0.164
-0.525
-0.575
-0.376
-0.359
-0.550
-0.709
-0.425
0.155
0.140
0.165
0.158
0.156
0.174
0.144
0.154
0.802
0.810
0.809
0.814
0.812
0.805
0.611
0.789
SPMN
SPMN
SPMN
SUM
SPMN
M
M
M
M
M
2.040
3.370
1.547
1.950
1.448
2.333
1.497
4.034
3.049
4.140
-0.250
0.058
-0.213
0.077
-0.164
0.146
0.192
0.097
0.025
0.084
-0.083
-0.422
-0.098
-0.039
0.175
0.154
0.200
0.132
0.122
0.129
0.786
0.809
0.792
0.809
0.797
Repository Habitat
!
0.287
0.189
0.216
0.199
0.209
0.168
0.189
0.419
0.348
"
-0.076
-0.787
-0.449
-0.491
-0.647
-0.241
-0.719
-0.109
-0.923
functional parameters
1/#
E
0.162
0.812
0.170
0.792
0.195
0.809
0.146
0.717
0.174
0.799
0.157
0.808
0.152
0.809
0.169
0.820
0.205
0.769
Species
Scaphella dubia
Voluta ebraea
Volutocorbis abyssicola
Olividae
Alcospira marginata
Amalda rubiginosa rubiginosa
Oliva porphyria
Oliva textilina
Olivella biplicata
Harpidae
Morum grande
Harpa major
Mitridae
Cancilla isabella
Mitra eremitarum
Mitra mitra
Mitra swainsoni
Costellariidae
Vexillum taeniatum
Pusia cancellaroides
Cystiscidae
Cystiscus nanaoensis
Cancellaroidea
Cancellariidae
Cancellaria nodulifera
Cancellaria spengleriana
Conoidea
Conidae
Conus pulicarius
Conus stimpsoni
SPMN
SPMN
SPMN
M
M
M
W
1.781
1.654
1.802
model parameters
T
D
3.712
-0.005
2.295
-0.238
2.526
-0.098
UMUT
SPMN
SPMN
SPMN
UMUT
M
M
M
M
M
1.713
1.686
1.916
1.753
1.952
2.557
2.962
0.674
0.837
1.925
-0.063
0.055
-0.192
-0.275
-0.074
0.154
0.087
-0.016
0.007
0.050
-0.310
-0.011
0.056
0.090
-0.144
0.156
0.133
0.142
0.151
0.168
0.808
0.809
0.775
0.757
0.807
SPMN
SPMN
M
M
2.287
2.686
1.467
0.874
-0.038
0.020
0.141
0.157
-0.883
-0.735
0.197
0.208
0.796
0.807
SPMN
SPMN
UMUT
SPMN
M
M
M
M
1.383
1.509
1.526
1.321
4.816
3.666
3.769
5.959
-0.138
-0.103
-0.495
-0.209
0.146
0.154
0.145
0.028
-0.025
-0.156
0.274
0.023
0.134
0.143
0.131
0.117
0.804
0.806
0.722
0.789
SPMN
UMUT
M
M
1.299
1.455
5.068
2.277
-0.169
-0.176
-0.007
0.379
0.222
-0.405
0.127
0.172
0.792
0.807
UMUT
M
1.733
1.034
-0.321
0.078
-0.078
0.153
0.750
UMUT
UMUT
M
M
2.245
1.920
1.113
1.761
0.009
-0.121
0.462
0.365
-0.432
-0.687
0.209
0.206
0.809
0.797
SUM
SPMN
M
M
1.394
1.355
1.184
1.129
0.088
-0.028
0.262
0.340
0.000
-0.336
0.144
0.154
0.796
0.767
Repository Habitat
!
-0.005
0.083
0.101
"
-0.267
-0.046
-0.065
functional parameters
1/#
E
0.123
0.808
0.146
0.788
0.147
0.807
Species
Conus euetrios
Conus textile
Conus capitaneus
Conus tulipa
Turridae
Bathytoma luhdorfi
Thatcheria mirabilis
Turris babylonia
Turris grandis
Terebridae
Subula muscaria
Oxymeris crenulatus
Heterobranchia
Architectonicoidea
Architectonicidae
Architectonica perspectiva
Architectonica taylori
Phsilaxis radiatus
Pyramidelloidea
Pyramidellidae
Longchaeus canaliculatus
Otopleura mitralis
Pyramidella dolabrata
Opithobranchia
Cephalaspidea
Actenoidea
Acteonidae
Acteon tornatilis
Acteon tornatilis
Pupa strigosa
SPMN
SPMN
SPMN
SPMN
M
M
M
M
W
1.448
1.742
1.752
2.132
model parameters
T
D
0.773
-0.097
1.484
-0.226
0.467
-0.065
1.615
-0.097
UMUT
SUM
UMUT
UMUT
M
M
M
M
1.429
1.694
1.316
1.263
3.826
1.362
4.401
6.265
-0.168
0.087
-0.248
-0.202
-0.116
-0.322
0.028
0.173
-0.243
-0.084
0.135
-0.308
0.133
0.148
0.132
0.125
0.789
0.820
0.783
0.790
UMUT
UMUT
M
M
1.193
1.233
8.682
6.914
-0.256
-0.275
0.368
0.381
0.029
-0.042
0.105
0.117
0.788
0.786
UMUT
UMUT
UMUT
M
M
M
1.650
1.613
1.821
0.794
0.789
0.744
0.366
0.335
0.269
0.295
0.677
0.184
-0.134
-0.418
-0.184
0.139
0.153
0.131
0.821
0.818
0.819
UMUT
UMUT
UMUT
M
M
M
1.209
1.324
1.234
4.901
3.045
4.099
-0.096
-0.230
-0.138
0.217
0.594
-0.030
-0.455
-0.949
-0.264
0.142
0.175
0.146
0.817
0.769
0.808
UMUT
UMUT
UMUT
M
M
M
1.718
1.864
1.902
2.071
1.825
2.101
-0.070
-0.043
-0.192
0.005
-0.021
0.099
-0.126
0.124
-0.557
0.149
0.144
0.172
0.807
0.809
0.791
Repository Habitat
!
0.012
0.012
-0.019
0.064
"
-0.221
0.109
0.130
-0.152
functional parameters
1/#
E
0.146
0.753
0.148
0.785
0.149
0.775
0.158
0.805
Species
Scaphandridae
Scaphander lignaria
Haminoeoidea
Haminoeidae
Atys naucum
Bulloidea
Bullidae
Bulla vernicosa
Bulla vernicosa
Hydatinoidea
Hydatinidae
Hydatina amplustre
Hydatina physis
Ringiculoidea
Ringiculidae
Ringiculina kurodai
Pulmonata
Basommatophora
Chilinoidea
Chilinidae
Chilina parchappei
Lymnaeoidea
Lymnaeidae
Radix swinhoei
Radix swinhoei
Radix auricularia
Lymnaea stagnalis
Planorboidea
Planorbidae
Gyraulus chinensis
Repository Habitat
W
model parameters
T
D
!
"
functional parameters
1/#
E
UMUT
M
1.732
1.611
-0.255
0.109
0.390
0.147
0.781
UMUT
M
2.219
0.323
0.014
0.080
-0.019
0.141
0.798
SUM
UMUT
M
M
1.479
1.634
0.964
1.600
-0.347
-0.215
0.145
0.135
-0.422
-0.567
0.152
0.168
0.699
0.789
UMUT
UMUT
M
M
3.355
5.311
1.262
1.376
-0.113
-0.101
0.019
0.087
-0.370
-0.124
0.179
0.193
0.803
0.806
UMUT
M
1.844
2.154
-0.133
0.045
-0.275
0.156
0.801
UMUT
F
2.190
2.277
-0.257
0.396
-0.521
0.207
0.795
UMUT
UMUT
UMUT
SHIN
F
F
F
F
2.501
3.024
4.001
2.031
1.910
1.854
0.976
2.331
-0.136
-0.218
-0.153
-0.159
0.321
0.299
0.334
0.384
0.058
-0.209
-0.342
-0.253
0.181
0.224
0.265
0.196
0.808
0.802
0.805
0.806
UMUT
F
1.361
0.251
0.483
0.701
-0.391
0.115
0.794
Species
Planorbarius corneus
Physidae
Physa acuta
Euplumonata
Ellobioidea
Ellobiidae
Cassidula nucleus
Ellobium chinense
Melampus flavus
Pythia pantherina
Stylommatophora
Elasmograta
Succineoidea
Succineidae
Succinea lauta
Orthurethra
Enoidea
Enidae
Buliminus sinaiensis
Sigmurethra
Clausiloidea
Clausiliidae
Achatinella apexfulva (A. leucorraphe)
Achatinella bulimoides mistura
Achatinella mustelina
Auriculella aurieula
Auriculella montana
Luchuphaedusa inclyta
Megalophaedusa martensi
Partulina splendida
SPMN
F
W
1.236
model parameters
T
D
!
0.388
0.322
-0.493
SHIN
F
1.933
2.291
-0.305
0.223
-0.719
0.186
0.757
UMUT
UMUT
UMUT
SUM
L
L
L
L
1.641
1.514
1.619
1.381
1.080
2.461
1.311
1.867
-0.037
-0.138
0.051
-0.172
0.209
0.217
0.054
0.317
-0.749
-0.080
-0.600
-0.253
0.170
0.154
0.156
0.158
0.796
0.815
0.816
0.785
SHIN
L
3.908
1.762
-0.407
0.407
-0.414
0.218
0.784
SPMN
L
1.303
4.937
-0.031
0.412
-0.842
0.138
0.797
BPBM
BPBM
BPBM
BPBM
BPBM
SPMN
SHIN
BPBM
L
L
L
L
L
L
L
L
1.530
1.484
1.547
1.452
1.584
1.203
1.208
1.550
1.962
2.225
2.180
2.819
2.328
6.958
7.603
2.582
-0.019
-0.068
-0.162
-0.108
-0.239
0.143
-0.029
-0.138
0.588
0.462
0.523
0.448
0.391
-0.098
0.048
0.558
0.472
-0.176
-0.225
-0.518
-0.557
-1.326
-0.618
-0.359
0.141
0.169
0.182
0.176
0.186
0.096
0.111
0.192
0.821
0.821
0.814
0.811
0.799
0.766
0.803
0.806
Repository Habitat
"
-0.738
functional parameters
1/#
E
0.116
0.701
Species
Partulina virgulate
Pinguiphaedusa platydera
Orthaloidea
Orthalicidae
Liguus virgineus
Orthalicus undatus
Orthalicus undatus
Orthalicus undatus
Placostylus hargravesi
Cerionidae
Cerion incanum
Bulimulidae
Acavus phoenix
Auris bilabiata
Bostryx tumidulus
Luchuena reticulata variegata
Luchuena reticulata variegata
Mirus japonicus
Neopetraeus arboriferus
Zebrina cylindrica
Bulimuloidea
Odontostomidae
Spixia alvarezi
Cyclodontina inflata
Achatinoidea
Achatinidae
Achatina bicarinata
Subulinidae
Rumina decollata
Streptaxoidea
BPBM
SHIN
L
L
W
1.596
1.234
model parameters
T
D
!
2.416
-0.168
0.495
5.208
-0.024
0.014
SPMN
SPMN
SPMN
SPMN
SPMN
L
L
L
L
L
1.383
1.782
1.676
1.729
1.615
3.829
2.007
2.295
2.046
3.434
-0.062
-0.050
-0.141
-0.121
0.003
0.562
0.495
0.544
0.564
0.227
-0.304
-0.277
-0.328
-0.197
-0.324
0.162
0.193
0.200
0.198
0.145
0.808
0.808
0.805
0.809
0.809
UMUT
L
1.166
4.590
-0.080
0.090
-0.478
0.146
0.808
SPMN
SPMN
UMUT
UMUT
UMUT
UMUT
UMUT
UMUT
L
L
L
L
L
L
L
L
2.990
2.003
1.598
1.357
1.317
1.289
1.469
1.257
1.183
2.364
1.982
3.124
3.713
4.371
3.047
4.600
-0.203
-0.111
-0.058
-0.014
-0.061
0.011
-0.069
-0.099
0.753
-0.043
0.464
0.369
0.324
0.192
0.346
0.289
-0.034
-0.627
-0.472
-0.080
-0.515
-0.714
-0.502
-0.543
0.190
0.151
0.186
0.153
0.154
0.140
0.165
0.145
0.802
0.793
0.817
0.822
0.816
0.804
0.806
0.814
UMUT
UMUT
L
L
1.279
1.286
4.996
3.868
0.020
-0.067
0.219
0.337
-0.671
-0.828
0.132
0.154
0.806
0.804
SHIN
L
1.613
2.745
-0.028
0.228
-0.407
0.158
0.808
SPMN
L
1.164
6.439
-0.114
0.285
-0.369
0.132
0.820
Repository Habitat
"
-0.141
-1.052
functional parameters
1/#
E
0.179
0.815
0.131
0.778
Species
Streptaxidiae
Euglandia rosea
Rhytidoidea
Rhytididae
Rhytida dunniae
Rhytidoidea
Haplotrematidae
Pedinogyra hayii
Acavoidea
Acavidae
Acavus haemastoma
Acavus haemastoma
Brazieresta naturalistarum
Caryodidae
Pedinogyra hayii
Zonitoidea
Zonitidae
Coxia macgregori
Oxychilus draparnaudi
Helicarionoidea
Helicarionidae
Ryssota lamarckiana
Ariophantidae
Megaustenia siamensis
Helicoidea
Helicidae
Eobania vermiculata
Helix pomatia
Cepaea hortensis
Cepaea nemoralis
Repository Habitat
W
model parameters
T
D
!
"
functional parameters
1/#
E
UMUT
L
1.812
3.092
-0.085
0.275
-0.238
0.160
0.808
UMUT
L
2.592
0.546
0.089
0.605
-0.312
0.195
0.814
SPMN
L
1.509
1.606
0.297
0.959
-0.070
0.164
0.809
SHIN
SHIN
UMUT
L
L
L
2.283
2.685
1.696
1.440
1.356
2.596
-0.054
-0.013
-0.075
0.879
0.869
0.334
-0.372
-0.262
-0.321
0.162
0.181
0.171
0.807
0.808
0.808
SHIN
L
1.436
0.636
0.240
1.059
-0.125
0.155
0.786
SPMN
UMUT
L
L
1.145
1.819
0.200
0.602
0.546
0.088
0.883
0.543
-0.100
-0.438
0.094
0.169
0.695
0.804
SPMN
L
2.662
0.632
-0.077
0.801
-0.209
0.235
0.810
SHIN
L
5.306
0.878
0.011
0.638
0.083
0.189
0.808
SPMN
SHIN
SHIN
SHIN
L
L
L
L
1.890
2.149
1.827
2.614
0.936
1.248
1.085
0.593
-0.190
-0.142
-0.075
0.126
0.926
0.524
0.868
0.512
-0.191
-0.492
-0.276
-0.407
0.191
0.232
0.227
0.197
0.781
0.807
0.811
0.810
Species
Bradybaenidae
Acusta despecta despecta
Aegista elegantissima
Aegista elegantissima
Aegista hatakedai
Aegista oculus
Bradybaena pellucida
Bradybaena similaris
Bradybaena similaris
Calocochlia zonifera
Chloraea stenopsis
Chrysallis caniceps
Chrysallis virgata
Chrysallis virgata
Chrysallis virgata
Euhadra aomoriensis
Euhadra aomoriensis
Euhadra aomoriensis
Euhadra aomoriensis
Euhadra herklotsi herklotsi
Euhadra quaesita
Euhadra quaesita
Euhadra quaesita
Euhadra sandai communis
Euhadra scaebola
Helicostyla marinduquensis
Helicostyla mirabilis
Trishoplita pallens
Camaenidae
Amphidromus atricallosus
Repository Habitat
W
model parameters
T
D
!
"
functional parameters
1/#
E
UMUT
UMUT
UMUT
SHIN
UMUT
SHIN
SHIN
SHIN
SPMN
UMUT
SHIN
SHIN
SHIN
SPMN
SHIN
SHIN
SHIN
SHIN
SPMN
SHIN
SHIN
SHIN
SPMN
SPMN
SPMN
SPMN
SHIN
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
1.862
1.661
1.717
1.747
1.453
2.123
1.791
1.778
1.687
1.812
1.466
1.573
1.392
1.454
1.941
1.910
1.868
1.990
1.851
1.802
2.087
2.158
2.007
1.716
1.767
2.284
1.774
1.099
0.495
0.549
0.681
0.774
0.900
0.872
0.882
1.688
0.878
3.427
2.973
3.815
3.607
0.691
0.827
0.739
0.808
0.805
1.041
0.814
0.550
0.664
0.563
1.852
1.214
0.693
-0.072
0.222
0.157
0.161
0.327
0.005
-0.140
0.058
-0.134
-0.043
-0.150
-0.152
-0.329
-0.406
0.048
0.075
0.056
0.057
0.089
0.028
0.067
0.102
0.096
0.162
-0.281
-0.143
0.117
0.515
0.956
1.023
0.798
1.032
0.637
0.617
0.645
0.724
0.679
0.389
0.559
0.482
0.508
0.662
0.644
0.696
0.655
0.839
0.460
0.705
0.417
0.803
0.994
0.687
0.715
0.671
-0.296
-0.021
-0.276
-0.033
-0.264
0.013
-0.160
0.112
-0.307
-0.461
-0.307
-0.536
-0.106
-0.531
-0.145
-0.075
-0.236
-0.423
-0.391
-0.581
-0.454
-0.317
-0.331
-0.213
-0.035
-0.242
-0.116
0.187
0.148
0.184
0.156
0.161
0.176
0.168
0.154
0.217
0.200
0.164
0.194
0.147
0.160
0.171
0.167
0.178
0.200
0.207
0.187
0.216
0.164
0.198
0.177
0.183
0.258
0.158
0.812
0.803
0.805
0.811
0.812
0.816
0.787
0.812
0.812
0.804
0.805
0.802
0.776
0.721
0.809
0.814
0.811
0.814
0.813
0.814
0.812
0.812
0.813
0.805
0.781
0.805
0.809
SHIN
L
1.585
2.616
-0.277
0.446
-0.432
0.183
0.787
Species
Amphidromus perversus
Amphidromus perversus
Helicostyla horida
Papuia hindei
Papuina chapmani
Papuina malantanensis
Papuina melanesia
Phoenicobius aratus
Pleurodonte lamarckii
Satsuma ferruginea
Satsuma japonica
Satsuma largillierti
Spaherospira appendiculata
Sphaerospira mattea
Sphaerospira mattea
Xanthomelon pachystylum
Helminthoglyptidae
Micrarionta kelletti
Hygromiidae
Helicigona faustina
SHIN
SHIN
SHIN
SPMN
SPMN
SPMN
SPMN
SHIN
UMUT
SHIN
SHIN
UMUT
SHIN
SHIN
SHIN
SPMN
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
L
W
1.496
1.645
1.595
1.591
1.764
1.633
1.822
1.435
1.615
1.648
1.555
1.637
1.798
1.624
1.699
2.233
model parameters
T
D
2.724
-0.115
2.429
-0.274
2.235
-0.215
2.895
-0.272
1.857
-0.198
1.703
-0.131
1.405
0.043
2.643
-0.077
0.819
-0.194
1.082
-0.052
1.327
-0.112
1.252
-0.143
1.280
-0.102
1.210
-0.133
0.889
-0.089
1.163
-0.140
UMUT
L
1.946
0.858
-0.082
0.847
-0.698
0.229
0.794
SPMN
L
1.832
0.637
0.145
0.852
-0.244
0.179
0.812
Repository Habitat
!
0.422
0.441
0.790
0.650
0.901
1.043
0.955
0.651
1.075
0.770
0.938
0.699
0.910
0.860
0.923
0.765
"
-0.280
-0.533
0.299
0.144
0.088
0.171
-0.306
-0.598
-0.085
-0.026
-0.327
-0.403
-0.527
-0.477
-0.360
-0.074
functional parameters
1/#
E
0.172
0.815
0.194
0.789
0.155
0.795
0.163
0.788
0.190
0.800
0.176
0.807
0.172
0.808
0.193
0.808
0.164
0.749
0.169
0.808
0.212
0.796
0.200
0.798
0.185
0.801
0.222
0.797
0.215
0.796
0.223
0.809